Selected Publications (2000-2013)


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  • SACCHI R., PELLITTERI-ROSA D, MARCHESI M., GALEOTTI P. & FASOLA M. 2013 A comparison among sexual signals in courtship of European tortoises. Journal of Herpetology 47: 215-221. I.F. 1.077.
  • GALEOTTI P., SACCHI R., PELLITTERI-ROSA D., BELLATI A., COCCA W., GENTILLI A., SCALI S. 2013. Colour Polymorphism and Alternative Breeding Strategies: Effects of Parent’s Colour Morph on Fitness Traits in the Common Wall Lizard. Evolutionary Biology 40: 385-394. DOI:10.1007/s11692-012-9222-3. I.F. 3. 61
  • SACCHI R., PELLITTERI-ROSA D, GALEOTTI P. & FASOLA M. 2013. Colour variation in the polymorphic common wall lizard (Podarcis muralis): an analysis using the RGB colour system. Zoologischer Anzeiger 252: 431-439. I.F. 


  • GALEOTTI P., BERNINI G., LOCATELLO L., SACCHI R., RUBOLINI D., FASOLA M. 2012. Sperm traits negatively covaries with size and asymmetry of a secondary sexual trait in a freshwater crayfish. PLOS/One 7(8). e43771. I.F. 4.092.doi:10.1371/journal pone.0043771.
  • SERRA L., PIRRELLO S., CAPRIOLI M., GRIGGIO M., ANDREOTTI A., ROMANO A., PILASTRO A., SAINO N., SACCHI R., GALEOTTI P., FASOLA M., SPINA F., RUBOLINI D. 2012. Seasonal decline of offspring quality in the European starling Sturnus vulgaris: an immune challenge experiment. Behavioural Ecology & Sociobiology 66: 697-709. I.F. 3.179
  • SACCHI R., PELLITTERI-ROSA D, GALEOTTI P., ZUFFI, M. A. L., FASOLA M. 2012. Studying the reproductive biology of the Common wall lizard Podarcis muralis using ultrasonography. Journal of Zoology 287: 301-310. I.F. 2.043.
  • PELLITTERI-ROSA D., SACCHI R., PUPIN F., BELLATI A., COCCA W., GENTILLI A., GALEOTTI P., FASOLA M. 2012. Testing the ability to store sperm: an experimental manipulation of mating opportunities in the common wall lizard, Podarcis muralis . Acta Herpetologica 7: 111-118. I.F. 0.580.
  • SACCHI R., PELLITTERI-ROSA D.,BELLATI A., SCALI S., TRONCONI D., GALEOTTI P., FASOLA M. 2012. Biological correlates of two dorsal color pattern types in the Common Wall Lizard, Podarcis muralis (Laurenti, 1768). Herpetozoa25: 3-11. I.F. 0.389.
  • DE MARCHI G., FASOLA M., CHIOZZI G., BELLATI A., GALEOTTI P. 2012. Sex discrimination of Crab Plovers (Dromas ardeola) by morphometric traits. Waterbirds 35: 332-337. I.F. 0.757.


  • BELLATI A., PELLITTERI-ROSA D., SACCHI R., NISTRI A., GALIMBERTI A., CASIRAGHI M., FASOLA M., GALEOTTI P. 2011. Molecular survey of morphological subspecies reveals new mitochondrial lineages in Podarcis muralis (Squamata: Lacertidae) from the Tuscan Archipelago (Italy). Journal of Zoological Systematics and Evolutionary Research 49: 240-250. I.F. 1.384
  • PELLITTERI ROSA D., SACCHI R., GALEOTTI P., MARCHESI M., FASOLA M. 2011. Courtship displays are condition-dependent signals that reliably reflect male quality in Greek Tortoises. Chelonian Conservation and Biology 10: 10-17. I.F. 0.808
  • GALEOTTI P., SACCHI R., PELLITTERI ROSA D., FASOLA M. 2011. The yellow cheek-patches of the Hermann’s tortoise: sexual dimorphism and relationship with body condition. Italian Journal of Zoology 78: 464-470. I.F. 0.843


  • Pellitteri-Rosa D., Sacchi R, Galeotti P., Marchesi m., Fasola M. 2010. Do Hermann’s tortoises (Testudo hermanni) discriminate colours? An experiment with natural and artificial stimuli. Italian Journal of Zoology 76: 481-491. I.F. 0.843
  • SACCHI R., SCALI S., PELLITTERI-ROSA D., PUPIN F., GENTILLI A., TETTAMANTI S., CAVIGIOLI L., RACINA L., MAIOCCHI V., Galeotti P., FASOLA M. 2010. Photographic identification in reptiles: a matter of scales. Amphibia Reptilia 31: 489-502. I.F. 0.976
  • Galeotti P., PELLITTERI-ROSA D., SACCHI R., GENTILLI A., PUPIN F., RUBOLINI D., FASOLA M. 2010. Sex-, morph- and size-specific susceptibility to stress measured by haematological variables in captive common wall lizard Podarcis muralis.Comparative Biochemistry & Physiology A. 157: 354-363. I.F. 2.134


  • GALEOTTI P., RUBOLINI D., PUPIN F., SACCHI R., ALTOBELLI E., NARDI P. A., FASOLA M. 2009. Presence of rivals reduces mating probability but does not affect ejaculate size in the freshwater crayfish Austropotamobius italicus.Behaviour 146: 45-68. I.F. 1.604
  • SACCHI R., PUPIN F. GENTILLI A., RUBOLINI D., SCALI S, FASOLA M., GALEOTTI P. 2009. Male-male combats in a polymorphic lizard: residency and size, but not colour, affect fighting rules and contest outcome.Aggressive Behavior 35: 274-283. I.F. 1.240
  • GALEOTTI P., RUBOLINI D., SACCHI R., FASOLA M. 2009. Global changes and animal phenotypic responses: melanin-based plumage redness of scops owls increased with temperature and rainfall during the last century.Biology Letters 5: 532-534. I.F. 3.521


  • RuBolini D., Brichetti P., Galeotti P., Fasola M. 2008. Recent decline of Italian Sparrow Passer italiae urban populations in northern Italy. IBIS 150: 177-181. I.F. 1.219
  • GALEOTTI P., RUBOLINI D., PUPIN F., SACCHI R., FASOLA M. 2008 Sperm removal and ejaculate size correlate with chelae asymmetry in a freshwater crayfish species. Behavioral Ecology and Sociobiology 62: 1739-1745. I.F. 2.917 


  • GALEOTTI P., PUPIN F., RUBOLINI D., SACCHI R., NARDI P. A., FASOLA M. 2007. Effects of female mating status on copulation behaviour and sperm expenditure in the freshwater crayfish Austropotamobius italicus. Behavioral Ecology and Sociobiology61: 711-718. I.F. 2.754
  • Ficetola G. F., Sacchi R., Scali F., Gentilli A. , De Bernardi, Galeotti P. 2007.Vertebrates respond differently to human disturbance: implications for the use of a focal species approach.Acta Oecologica 31: 109-118. I.F. 1.306.
  • Rubolini D., Sacchi R., Pupin F., Scali F., Gentilli A., Razzetti E, Galeotti P., Fasola M. 2007. Morph-specific immunity in male Podarcismuralis. Amphibia-Reptilia 28: 408-412. I.F. 0.929
  • Sacchi R, Scali S, Galeotti P., Fasola M 2007. The numerical encoding of scale morphology highly improves photographic identification in lizards. Acta Herpetologica2: 27-35. I.F. 0.580.
  • RUBOLINI D., GALEOTTI P., PUPIN F., SACCHI R., FASOLA M: 2007. Repeated mating and sperm depletion in relation to body size among freshwater crayfish Austropotamobius italicus males. Freshwater Biology 52: 1898-1906 I.F. 2.650.
  • Sacchi R., Pupin F., Zuffi M. A. L., Scali S., Boncompagni E., Binda A., Galeotti P., Fasola M. 2007. Blood cell morphology of the Moorish gecko, Tarentola mauritanica. Amphibia-Reptilia 28: 503-508.I.F. 0.929.
  • GALEOTTI P., SACCHI R., PELLITTERI ROSA D., FASOLA M. 2007. Olfactory discrimination of species, sex, and sexual maturity by the Hermann’s tortoise Testudo hermanni. Copeia 2007 (4): 980-985. I.F. 1.117.
  • GALEOTTI P., RUBOLINI D. 2007. Head ornaments in owls: what are their functions? Journal of Avian Biology 38: 731-736 I.F. 2.381.
  • Sacchi R., Scali S., Pupin F., Gentilli A., Galeotti P., Fasola M. 2007. Microgeographic variation of colour morph frequency and biometry of common wall lizards. Journal of Zoology 273: 389-396. I.F. 1.516.
  • PUPIN F., SACCHI R., GENTILLI A., GALEOTTI P., FASOLA M. 2007. Discrimination of 
  • toad calls by smooth newts: support for the heterospecific attraction hypothesis. Animal Behaviour 74:1683-1690. I.F. 2.752


  • GHERARDI F., RENAI B., GALEOTTI P., RUBOLINI D. 2006. Nonrandom mating, mate choice, and male-male competition in the crayfish Austropotamobius italicus, a threatened species. Archive für Hydrobiologie165: 557-576. I.F. 1.362
  • GALEOTTI P., RUBOLINI D., FEA G., GHIA D., NARDI P. A.,GHERARDI F., FASOLA M. 2006. Female freshwater crayfish adjust egg and clutch size in relation to multiple male traits. Proceedings of the Royal Society of London B273: 1105-110. I.F. 3.612
  • RUBOLINI D., GALEOTTI P., FERRARI G., SPAIRANI M., BERNINI F., FASOLA M. 2006. Sperm allocation in relation to female size, male traits and copulation behaviour in a freshwater crayfish. Behavioral Ecology and Sociobiology60: 212-219. I.F. 2.316
  • RUBOLINI D., PUPIN F., SACCHI R., GENTILLI A., ZUFFI M.A.L., GALEOTTI P., SAINO N. 2006. Sexual dimorphism in digit length ratios in two lizard species. Anatomical Records A288A: 491-497. I.F. 1.973


  • GALEOTTI P., SACCHI R., PELLITTERI ROSA D., FASOLA M. 2005. Female preference for fast-rate, high-pitched calls in Hermann’s Tortoises Testudo Hermanni. Behavioral Ecology16: 301-308. I.F. 2.943
  • GARAMSZEGI L.Z., BALSBY T.J.S, BELL B.D., BOROWIEC M., BYERS B. E. DRAGANOIU T., EENS M., FORSTMEIER W., GALEOTTI P, GIL D., GORISSEN L., HANSEN P., LAMPE H. M., LEITNER S., LONTKOWSKI J., NAGLE L., NEMETH E., PINXTEN R., ROSSI J-M., SAINO N., TANVEZ A., TITUS R., TÖRÖK J., VAN DUYSE E., MØLLER A. P. 2005. Estimating the complexity of bird song by using capture-recapture approaches from community ecology. Behavioral Ecology and Sociobiology57: 305-317. I.F. 2.232
  • RUBOLINI D., ROMANO M., BONCORAGLIO G., FERRARI R. P., MARTINELLI R., GALEOTTI P., FASOLA M., SAINO N. 2005. Effects of elevated egg corticosterone levels on behavior, growth and immunity of yellow-legged gull (Larus michahellis) chicks. Hormones & Behavior47: 592-605. I.F. 3.737
  • GALEOTTI P., SACCHI R., FASOLA M., BALLASINA D. 2005. Do mounting vocalisations in tortoises have a communication function? A comparative analysis. Herpetological Journal15: 61-71. I.F. 0.706
  • GALEOTTI P., SACCHI R., VICARIO V. 2005. Fluctuating asymmetry in body traits increases predatrion risks: tawny owl selection against asymmetric woodmice. Evolutionary Ecology19: 405-418. I.F.1.776
  • GALEOTTI P., SACCHI R., FASOLA M., PELLITTERI ROSA D., MARCHESI M., BALLASINA D. 2005. Courtship displays and mounting calls are honest, condition-dependent signals influencing mating success in Hermann’s tortoises. Canadian Journal of Zoology83: 1306-1313. I.F. 1.175


  • FASOLA M., GALEOTTI P., NIANHUA DAI, YUANHUA DONG, YNGMEI ZHANG 2004. Large numbers of breeding egrets and herons in China. Waterbirds27: 126-128. I.F. 0.381
  • GALEOTTI P., RUBOLINI D. 2004. The niche variation hypothesis and the evolution of colour polymorphism in birds: a comparative study of owls, nightjars and raptors. Biological Journal of Linnean Society82: 237-248. I.F. 1.935
  • SACCHI R., GALEOTTI P.,FASOLA M., GERZELI G. 2004. Larinx morphology and its relationship with sound production in three species of Testudinidae. Journal of Morphology261: 175-183. I.F. 1.528


  • SACCHI R., GALEOTTI P., FASOLA M., BALLASINA D. 2003 Vocalizations and courtship intensity correlate with mounting success in marginated tortoises Testudo marginata. Behavioral Ecology Sociobiology55: 95-102. 2.649
  • GALEOTTI P., SACCHI R. 2003. Differential parasitemia in the tawny owl (Strix aluco): effects of colour morph and habitat. Journal of Zoology261: 91-99. I.F. 1.175
  • GALEOTTI P., RUBOLINI D., DUNN P. O., FASOLA M. 2003. Colour polymorphism in birds: causes and functions. Journal of Evolutionary Biology. 16: 635-646. I.F. 3.010
  • SAINO N., GALEOTTI P., SACCHI R., BONCORAGLIO G., MARTINELLI R., MOLLER A. P. 2003. Sex differences in begging vocalisations of nestling barn swallows (Hirundo rustica). Animal Behaviour66: 1003-1010. I.F. 2.557
  • SAINO N., ROMANO M., SACCHI R., NINNI P., GALEOTTI P., MOLLER A. P. 2003. Do male barn swallows (Hirundo rustica) experience a trade-off between the expression of multiple sexual signals? Behavioral Ecology and Sociobiology54: 465-471. 2.649


  • SACCHI R., SAINO N., GALEOTTI P. 2002. Features of begging calls reveal general condition and need of food of barn swallow (Hirundo rustica) nestlings. Behavioral Ecology13: 268-273. I.F. 2.449
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